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Definition of "Aves"?
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How should "Aves" be defined?
As a crown group (e.g., sensu Gauthier 1986)
23%
 23%  [ 3 ]
As a total group (e.g., sensu Ax)
0%
 0%  [ 0 ]
As a non-crown node-based clade (e.g., sensu Chiappe 1997, Sereno 1997)
30%
 30%  [ 4 ]
As a non-total branch-based clade (e.g., sensu Marjanovic [unpublished])
30%
 30%  [ 4 ]
As an apomorphy-based clade (please specify).
15%
 15%  [ 2 ]
Other (please explain).
0%
 0%  [ 0 ]
Total Votes : 13

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ottscay



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PostPosted: Fri 08 Dec, 2006 2:39 am    Post subject: Reply with quote

That doesn't really bother me, but aren't we then asking neontologists (and specifically ornithologists) to abondon "Aves" altogether? And what would we call a "bird"? I realize that bird isn't a phylogenetic term, but pretending like the term "bird" doesn't have to be connected to our cladograms is just shoving the task onto educators and science popularizers.
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tmkeesey
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PostPosted: Fri 08 Dec, 2006 2:51 am    Post subject: Reply with quote

(I'm totally playing devil's advocate here, just to be clear--I doubt leaving the name undefined would be a satisfactory answer to the problem.)

Not all vernacular terms, even those that are monophyletic in most usages, need to correspond directly to a clade name. Consider "human". Does it correspond to Homo sapiens? To Homo? To Hominina? Is Homo habilis human? What about Paranthropus? Australopithecus? Praeanthropus? It's a vague term, at least in paleontology, even though under all usages it is monophyletic.
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ottscay



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PostPosted: Fri 08 Dec, 2006 9:17 pm    Post subject: Reply with quote

"Human", of course, is usually used as more of an ontological statement of the recipients' worth in a moral or ethical sense; the problem historically is usually with people not extending the term far enough (e.g. to other members of H. sapiens) rather than too far, although your points are logically valid.

Personally I'd restrict it to our species, until such time as we can more accurately evaluate the cognitive abilities of extinct hominids (after all, it's hard to abuse the civil rights of extinct organisms...), but physical anthropologist Wolpoff would disagree with my "limited" nomenclatural distribution.

In practice, I think the additional qualatative socio/political aspects attributed to the term "human" makes it non-equivalent to other vernacular terms, such as fish, lizard, or bird (or Aves).
But hey, what do others thing?
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tmkeesey
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PostPosted: Fri 08 Dec, 2006 9:49 pm    Post subject: Reply with quote

"Human" has some other baggage, sure, but it has the exact same problem as "bird". Some other placentalian examples:

Does "chimp" apply to Pan troglodytes? Pan? Basal (hypothetical) members of the Pan lineage, after separating from Hominina?

Does "horse" apply to Equus caballus? Equus? Equinae? Equidae? Is Eohippus a horse? What about Merychippus or Pliohippus?

Does "whale" apply to the crown group? The total group? Some apomorphy-based clade in between? Is Ambulocetus a whale? Pakicetus?

So on and so on.
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dinoguy2



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PostPosted: Sat 09 Dec, 2006 3:04 pm    Post subject: Reply with quote

ottscay wrote:

Shoe-horning the great diversity of feathered terrestrial maniraptorans into being birds not only could blind us to their own unique adaptaions, but it stops us from examing where basal birds truly aaquired other avian characters. Both Velociraptors and Archaeopteryx are more like basal archosaurs than modern birds in their ability to stablize their shoulder joint during wing excursion; this may have sever implications for the supposed flight ability of Archaeopteryx.


Why not just abandon the term bird altogether? Surely Sinornis is closer to basal archosaurs than hummingbirds are, so doesn't referring to enantiornithines as birds create the same problem? Obviously, when dealing with a gradiant like evolution, assigning names to anything is always going to obscure something, or add bias to something, depending on context.

Quote:

I am frankly ok with restricting "bird" and "aves" to the crown group, I just don't think it's the only viable solution (e.g. David's excellent suggestion). But it seems entirely misplaced to decide that all feathered dinosaurs are birds, simply beause we're looking at the question in a very macroscopic "they're more birds than reptiles" sort of way.


Why shouldn't the macrosopic view be an option? Why not be able to look at it from a wide-angle, bird or reptile typological view in one context and be able to look at it in a specific, morphological "well, it's clocer to the MRCA of Aviale than to derived Neornithes" when getting down to the nitty-gritty?

""Human" has some other baggage, sure, but it has the exact same problem as "bird". Some other placentalian examples:"

I had written up a whle paragraph using this example, but TMK said it better. Let vernacular terms be what they are--semi-vague typological distinctions. it's a whale if it looks more like a wahle than a basal artiodactyl (or whatever it is this week Wink). It's a human if it looks more like me than a chimp (a fine distinction). It's a bird if it looks more like a bird than a crocodile. These are all vague, subjective "definitions" (let's figure out which clade "fish" or "lizard" correspond to!). If you want to be specific, simply do not use the term "bird" in a scientific context. Use Avian, figure out a consensus on what "Avian' means without regard to what a "bird" is, and let bird remain "more like a bird than like a reptile".

Anyway, Aves as currently used by most is just a name on a cladogram. It might as well be replaced with a number. The strong desire that it correspond with "bird" strikes me as almost hypocritical from people who despises the old Linnean system so much. What if phylocode is published and Aves is defined as Passer + Struthio? Wouth everyone here be ok with the fact that Ichthyornis is a non-avian dinosaur? Would you still call it a bird? Why?

And what if, subsequently, Apteryx turns out to be non-avian?
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Nescio



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PostPosted: Sat 09 Dec, 2006 9:37 pm    Post subject: Reply with quote

Quote:
It's a bird if it looks more like a bird than a crocodile.

That'd make Pteranodon a bird ... are you calling for a total group definition? Wink

More seriously, I think defining Aves as Struthio + Passer would be stupid, because that clade is either = Neornithes or smaller than Neornithes. Most people who'll use these terms are erudite enough to recognize the stems, and having Birds be a subgroup of New Birds is potentially confusing as well as silly.



(As for fish, I tend to equate it with Craniata. Among other advantages, this allows me to insist that whales are fish. Razz )
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Eoraptor



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PostPosted: Sun 10 Dec, 2006 12:35 am    Post subject: Reply with quote

I've generally assumed total groups were used with mammalian vernacular taxa. So yes Pakicetus is a whale (as long as dolphins and porpoises are types of whales), Eohippus is a horse, and all Pan-Pan taxa are chimpanzees. I'm sure it would fail in some cases though...
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dinoguy2



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PostPosted: Sun 10 Dec, 2006 4:59 pm    Post subject: Reply with quote

Eoraptor wrote:
I've generally assumed total groups were used with mammalian vernacular taxa. So yes Pakicetus is a whale (as long as dolphins and porpoises are types of whales), Eohippus is a horse, and all Pan-Pan taxa are chimpanzees. I'm sure it would fail in some cases though...


Equating total groups with vernacular names could probably work in a lot of instances, though I guess only George Olshevsky would be happy with Brachiosaurus as a "bird"... Unless we come up with vernacular terms for sauropods, ornithischians, etc, and then let "bird" refer to Theropoda? Wink
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tmkeesey
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PostPosted: Sun 10 Dec, 2006 6:36 pm    Post subject: Reply with quote

Eoraptor wrote:
I've generally assumed total groups were used with mammalian vernacular taxa. So yes Pakicetus is a whale (as long as dolphins and porpoises are types of whales), Eohippus is a horse, and all Pan-Pan taxa are chimpanzees. I'm sure it would fail in some cases though...


I think it fails with Ambulocetus. I seriously doubt any layperson would identify this as a "whale". It would probably also fail with "bat", since there should be some shrew-like, utterly wingless stem-chiropterans (although such fossils are not known, or at least not recognized).

I've done this test with a few laypeople. Typically if shown a picture of, e.g., Velociraptor and asked if it's a bird, the response is, "Kind of...," or "Well, it's birdlike...."
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tmkeesey
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PostPosted: Sun 10 Dec, 2006 6:40 pm    Post subject: Reply with quote

dinoguy2 wrote:
Equating total groups with vernacular names could probably work in a lot of instances, though I guess only George Olshevsky would be happy with Brachiosaurus as a "bird"... Unless we come up with vernacular terms for sauropods, ornithischians, etc, and then let "bird" refer to Theropoda? ;)


Ugh.

I would have to agree with Gauthier that generally, if anything, the crown group is closer to the vernacular meaning. Extending such terms beyond the crown group leads to unjustified inferences.

Edit: I can't recall if Gauthier's made that point about vernacular terms, but he certainly has made it about clade names.
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leowsham



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PostPosted: Mon 11 Dec, 2006 2:53 am    Post subject: Vernacular names Reply with quote

tmkeesey wrote:
I would have to agree with Gauthier that generally, if anything, the crown group is closer to the vernacular meaning.

This is, naturally, owing to the fact that we only had extant lifeform to give vernacular names to, without knowledge or insight about fossils back then. But there's a twist: vernacular names usually convey or stick to a particular character of a group, and so now when we learnt of Earth's history and its heritage of fossils, we'd like to extend vernacular names down to include basal forms that (seem to) bear the same character.
This belies the logical fault that we have been arguing here: evolution is a continuum of change, but psychologically we wish there were a watershed to stick a vernacular to (so as to let it be meaningful, to the layman at least).
My personal opinion is not to define "bird" (i.e. using "birdiness" to define Aves), but give a node or branch based definition to Aves (see previous arguments on Archaeopteryx, above) - let the laymen perceive where the ash shall fall!
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David Marjanović



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PostPosted: Tue 12 Dec, 2006 6:28 pm    Post subject: Reply with quote

Well, erm.

1) Leaving anything undefined is not really an option under the PhyloCode: if anyone wants to define it and gets it through peer review after "January 1, 200n", it is defined. Though maybe we should have a List of Trashed Names – after all, the ICZN has one (OK, "Rejected Names").

2) Treating "birds" and Aves as somehow separate won't work, because the Latin – and Spanish, and Portuguese – word "aves" means "birds". You will never manage to get the two apart. We might barely manage that trick with "whales", but not with "birds".

3)
"The feathers don't make the bird. The bird makes the feathers."
– Jacques Gauthier, at the first PhyloCode meeting
So there, just to have said it. Smile Actually, Jacques was paraphrasing Linnaeus, who wrote that the characters don't give the genus, the genus gives the characters, and this in spite of the fact that Linnaeus was an essentialist. (Darwin used that quote to good effect.)
Actually, there are a few cases where I think apomorphy-based definitions are both feasible and required by Article 10.1 (Tetrapoda, Testudinata sensu Lee 1995 and Joyce et al. 2004, Diapsida), but in general I can only recommend against them. The main reason is that, in order to tell whether something is part of a taxon the name of which has an apomorphy-based definition, it's not enough if we can establish its phylogenetic position, we also need to know if that particular character state was present. Try that with any kind of "feathers"...

4) Have I mentioned my Plan B yet? I like to forget it again and again. Smile If the crown-groupers get their wish to make Ichthyornis a nonavian dinosaur – and, of course, all our base are belong to them –, I'll simply take my definition, slap the name Ornithes on it, and publish that – just the same in Greek. While somewhat suboptimal, it would have the advantage of providing a more obvious etymological anchor for all existing the names that end in -ornithes, and it would nicely fit the tradition of using former synonyms for nested clades (Eutheria-Placentalia, Metatheria-Marsupialia, Ophidia-Serpentes, Proganosauria-Mesosauridae, Gymnophiona-Apoda, Caudata-Urodela, Salientia-Anura...).
George Olshevsky once said he wanted to have Aves for the crown-group and Ornithes for the total group. I don't know if he ever got that published.

5) Scott, why did you make the snout an exact copy of the PDW dromaeosaurids? I still haven't seen any evidence for the naked snout – contradicted by Eoenantiornis – or the "proto-beak" (probably also contradicted by that little birdie).
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tmkeesey
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PostPosted: Tue 12 Dec, 2006 6:47 pm    Post subject: Reply with quote

David Marjanović wrote:
4) Have I mentioned my Plan B yet? I like to forget it again and again. :-) If the crown-groupers get their wish to make Ichthyornis a nonavian dinosaur – and, of course, all our base are belong to them –, I'll simply take my definition, slap the name Ornithes on it, and publish that – just the same in Greek.


I really like that idea.
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Nescio



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PostPosted: Tue 12 Dec, 2006 11:23 pm    Post subject: Reply with quote

David Marjanović wrote:

Actually, there are a few cases where I think apomorphy-based definitions are both feasible and required by Article 10.1 (Tetrapoda, Testudinata sensu Lee 1995 and Joyce et al. 2004, Diapsida), but in general I can only recommend against them.

For what it may be worth, I'm inclined to agree with the Palaeos folks that an apomorphy-based definition won't work for Tetrapoda, what with all the weird stuff lurking between Eusthenopteron and Acanthostega these days. A definition strict enough to cut a line among these critters will end up makine poorly known taxa unclassifiable. But I suppose this discussion should be continued, if at all, in the Chordata forum.
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tmkeesey
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PostPosted: Wed 13 Dec, 2006 12:17 am    Post subject: Reply with quote

Yes, in fact, there's already a thread for it: Definition of "Tetrapoda"
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